NEGOMBO, Sri Lanka — A seaside Sri Lankan fishing town of Negombo has long been called “Little Rome,” a reference to its abundance of churches and its place at the center of the country’s small Catholic community.

But it was also known for its tolerance. Negombo’s Angurukaramulla Temple is a well-known stop for Buddhist pilgrims. Its Grand Mosque is famed for its beauty. The Hindu god Rama is said to have been nearby before his great battle with the demon-king Ravana.

During Sri Lanka’s long and bloody civil war, Negombo, about an hour north of the capital, Colombo, was largely spared the violence that raged elsewhere in the country.

“It is a safe haven for all,” Cardinal Malcolm Ranjith said at a religious conference held in the town in 2016.

But all that changed Sunday when a bomb ripped through Negombo’s St. Sebastian’s Church, one of a half-dozen coordinated attacks on churches and high-end hotels that killed nearly 300 people. At least 110 were killed in the attack at St. Sebastian’s, Ranjith said Monday.

“I was shocked when I saw the horrible devastation caused at the church,” he said after visiting St. Sebastian’s, pleading for tolerance and urging Christians not to seek revenge. Authorities have blamed the Sri Lankan Muslim group National Thowfeek Jamaath.

But in the hours after the blasts, even Ranjith called on officials to “mercilessly” punish those responsible for attacks, saying “only animals can behave like that.”

Catholicism is everywhere in Negombo, a town of about 140,000 people with dozens of churches and perhaps hundreds of small roadside Catholic shrines. About 65 percent of Negombo is Catholic, according to census data, though Catholics make up just 6 percent of the country’s population.

Negombo’s economy has long been dominated by fishing, but the tourism industry has grown quickly in recent years because of the town’s beaches and its proximity to the country’s international airport.

On Monday, many people came to pray in the garden outside the church, gathering at a statue of St. Sebastian, an early Christian martyr who was riddled with arrows during Roman persecutions.

Kumari Siriwardane had tears in her eyes as she spoke.

“I feel so sad about our village, our people, our religion,” said Siriwardane, 58. “The doors of our houses are closed.”

She struggled to find words to express her pain: “I feel very sad. Very very sad.”